Lewis, Christiane Nüsslein-Volhard 1993 Günter Blobel 1994 Stanley B. Prusiner 1995 Peter C. Doherty, Jack L. Strominger, Emil R. Unanue, Don C. Wiley, Rolf M. Zinkernagel 1996 Robert F. Furchgott, Ferid Murad 1997 Mark S. Ptashne 1998 Leland H. Hartwell, Yoshio Masui, Paul Nurse 1999 Clay Armstrong, Bertil Hille, Roderick MacKinnon 2000 Aaron Ciechanower, Awram Herszko, Alexander Varshavsky 2001 Mario Capecchi, Martin Evans, Oliver Smithies 2002 James E. Rothman, Randy W. Schekman 2003 Robert G. Roeder 2004 Pierre Chambon, Ronald M. Evans, Elwood V. Jensen 2005 Ernest McCulloch, James Till 2006 Elizabeth Blackburn, Carol W. Greider, Jack Szostak 2007 Ralph M. Steinman 2008 Victor R. Ambros, David C. Baulcombe, Gary B. Ruvkun 2009 John Gurdon, Shin’ya Yamanaka 2010 Douglas L. Coleman, Jeffrey M. Friedman 2011 Franz-Ulrich Hartl, Arthur L. Horwich 2012 Michael Sheetz, James Spudich, Ronald Vale 2013 Richard H. Scheller, Thomas C. SüdhofLasker Awards 2013 2014 Kazutoshi Mori, Peter WalterLasker Awards 2014 2015 Stephen J.