europeiske underslekter: Adelopterus Adelosia Aphaon Argutor Bothriopterus Calopterus Cheporus Cophosus Corax Cryobius Eosteropus Feronidius Haptoderus Haptotapinus Iberopus Lenapterus Lianoe Lyrothorax Melanius Metallophilus Morphnosoma Oreophilus Parahaptoderus Parapterostichus Petrophilus Phonias Platypterus Platysma Pseudomaseus Pseudorites Pseudosteropus Pterostichus Pyreneorites Sterocorax Steropus
Pterostichus er en slekt av billene i delgruppen (underfamilien) Pterostichinae i familiegruppen løpebiller. Systematikken følger Carabidae of the World.
Utseende Stor sett middelsstore (oftest 8-20 millimeter), avlange, blanke, mørkt brunlige til svarte løpebiller, noen ganger med en svak metallglans. Kjevene er ikke særlig forstørrede. På hodet er det to børster like innefor hvert fasettøye. Pronotum er mer eller mindre avrundet firkantet, gjerne med to grunne, grovt punkterte lengdefurer ved roten. Dekkvingene har fine punkstriper, ved ytterkanten nær spissen er kantlisten "vridd".
Levevis Artene er rovdyr som lever på bakken. Mange av dem er tallrike i skog eller på myrer, noen også på dyrket mark. De fleste er nattaktive.
Systematisk inndeling Pterostichus er en stor slekt med ca. 1200 kjente arter. Av og til kalles slekten Feronia, men det navnet betraktes oftest som et synonym for Poecilus.
ordenen biller, Coleoptera Linnaeus, 1758 underordenen Adephaga Schellenberg, 1806 overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802 familien løpebiller, Carabidae Latreille, 1802 underfamilie Pterostichinae Bonelli, 1810 slekten Pterostichus Bonelli, 1810 – nordeuropeiske arter: Pterostichus adstrictus Eschscholtz, 1823 Pterostichus aethiops (Panzer, 1796) Pterostichus anthracinus (Panzer, 1795) Pterostichus aterrimus (Herbst, 1784) Pterostichus crenatus (Duftschmid, 1812) Pterostichus cristatus (Dufour, 1820) Pterostichus diligens (Sturm, 1824) Pterostichus gracilis (Dejean, 1828) Pterostichus longicollis (Duftschmid, 1812) Pterostichus macer (Marsham, 1802) Pterostichus madidus (Fabricius, 1775) Pterostichus melanarius (Illiger, 1798) Pterostichus minor (Gyllenhal, 1827) Pterostichus niger (Schaller, 1783) Pterostichus nigrita (Paykull, 1790) Pterostichus oblongopunctatus (Fabricius, 1787) Pterostichus quadrifoveolatus Letzner, 1852 Pterostichus rhaeticus Heer, 1837 Pterostichus strenuus (Panzer, 1796) Pterostichus vernalis (Panzer, 1795) Liste over ikke-nordeuropeiske arter
Pterostichus aanistschenkoi O. Berlov & E. Berlov, 1999 Pterostichus aapsorum Belousov, 1991 Pterostichus abaciformis Straneo, 1955 Pterostichus abagonensis Reitter, 1896 Pterostichus abasarukini O. Berlov & E. Berlov, 1996 Pterostichus abasinus Belousov, 1991 Pterostichus abaxoides Dejean, 1828 Pterostichus abensis Morita, 2010 Pterostichus abishirensis Belousov, 1991 Pterostichus abnormis (J.Sahlberg, 1880) Pterostichus abrogatus Scudder, 1890 Pterostichus abudarini O. Berlov & E. Berlov, 1997 Pterostichus acrogonus (Chaudoir, 1843) Pterostichus acuspina (Tschitscherine, 1901) Pterostichus acutidens (Fairmaire, 1889) Pterostichus acutipes Barr, 1971 Pterostichus adatarasanus Sasakawa, 2005 Pterostichus ademidovae O.Berlov, 1996 Pterostichus adoxus (Say, 1823) Pterostichus adulterinus Wrase & J.Schmidt, 2006 Pterostichus aemiliae Facchini & Sciaky, 2000 Pterostichus aeneocupreus (Fairmaire, 1887) Pterostichus aenescens Chaudoir, 1850 Pterostichus aeneus Fischer von Waldheim, 1829 Pterostichus aereipennis (Solsky, 1871) Pterostichus agilis Allegro & Sciaky, 2010 Pterostichus agonus G.horn, 1880 Pterostichus aibgensis Starck, 1890 Pterostichus aimaki Jedlicka, 1968 Pterostichus ainus Plutenko, 2005 Pterostichus akitai Morita, 2004 Pterostichus akkusianus Kirschenhofer, 1981 Pterostichus akozyrevi O. Berlov & E. Berlov, 1999 Pterostichus aksekianus (Straneo, 1988) Pterostichus alacer A.Morawitz, 1862 Pterostichus alamedae (Casey, 1918) Pterostichus alexandrovi Lafer, 1979 Pterostichus alexeji Zamotajlov & Kryzhanovskij, 1992 Pterostichus algidus Leconte, 1852 Pterostichus altaianus Jedlicka, 1958 Pterostichus altaicus (Germar, 1824) Pterostichus altaiensis Poppius, 1906 Pterostichus amadeus Bousquet, 2012 Pterostichus amagisanus Tanaka & Ishida, 1972 Pterostichus amanoi Nakane, 1968 Pterostichus amaroides Dejean, 1828 Pterostichus ambigenus Bates, 1883 Pterostichus ambiguus (Fairmaire, 1858) Pterostichus amblypterus Chaudoir, 1868 Pterostichus amethystinus Mannerheim, 1843 Pterostichus amoenus Dejean, 1828 Pterostichus ampliatus Bates, 1882 Pterostichus amurensis Poppius, 1906 Pterostichus analis Jedlicka, 1963 Pterostichus anatolicus Jedlicka, 1963 Pterostichus andreae (Tschitscherine, 1897) Pterostichus andreinii Dodero, 1922 Pterostichus andrewesi Jedlicka, 1931 Pterostichus angoarnigi Morvan, 1994 Pterostichus angustus (Dejean, 1828) Pterostichus annamita (Straneo, 1939) Pterostichus annosus Casey, 1913 Pterostichus anomalus Chaudoir, 1878 Pterostichus anomostriatus Sciaky, 1995 Pterostichus antepunctatus Straneo, 1949 Pterostichus apenninus Dejean, 1831 Pterostichus apfelbecki Csiki, 1908 Pterostichus apiculatiphallus Nemoto, 1988 Pterostichus aralarensis Espanol & Mateu, 1945 Pterostichus araraticus Kirschenhofer, 1987 Pterostichus arator (Faldermann, 1836) Pterostichus arcanoides Lorens, 1998 Pterostichus arcanus Casey, 1913 Pterostichus archangaicus Shilenkov, 2000 Pterostichus arcticola (Chaudoir, 1868) Pterostichus arcuaticarinatus Kasahara, 1986 Pterostichus argutoriformis Poppius, 1906 Pterostichus aristochroides Deuve, 2006 Pterostichus armenus (Faldermann, 1836) Pterostichus arnoldii Erjiomin, 1996 Pterostichus arribasi Ortuño, 1991 Pterostichus arrowi Jedlicka, 1936 Pterostichus arrowianus Jedlicka, 1938 Pterostichus arsenjevi Lafer, 1979 Pterostichus arvernus Jeannel, 1937 Pterostichus asahinus Habu & Baba, 1960 Pterostichus asiaticus Poppius, 1906 Pterostichus asturicus (Jeanne, 1965) Pterostichus astutus (Tschitscherine, 1903) Pterostichus asymmetricus Bates, 1883 Pterostichus atratus Gistel, 1857 Pterostichus atrox (Andrewes, 1937) Pterostichus aubryi Coiffait, 1952 Pterostichus audax (Tschitscherine, 1895) Pterostichus auratus Heer, 1837 Pterostichus auriga Ball, 1962 Pterostichus avaricus Kryzhanovskij & Abdurakhmanov, 1983 Pterostichus aztecus Tschitscherine, 1897 Pterostichus badzhaliensis Sundukov, 2025Sundukov, Y.N. (2025) Three new species of the subgenus Cryobius Chaudoir, 1838 (Coleoptera: Carabidae, Pterostichus Bonelli, 1810) from the Russian Far East. Zootaxa, 5715 (1), 446–455. https://doi.org/10.11646/zootaxa.5715.1.39 Pterostichus baenningeri Jedlicka, 1931 Pterostichus balachowskyi Morvan, 1972 Pterostichus baldwini (Casey, 1924) Pterostichus balikliensis Hovorka & Skoupy, 2007 Pterostichus balthasari Jedlicka, 1937 Pterostichus balu Schmidt, 2009 Pterostichus bandotaro Tanaka, 1958 Pterostichus bannaticus Dejean, 1828 Pterostichus barbarae Sciaky & Facchini, 2003 Pterostichus barbarinus Casey, 1913 Pterostichus barbatus Fischer von Waldheim, 1829 Pterostichus bargusinicus Shilenkov, 2000 Pterostichus barlensis Straneo, 1935 Pterostichus barri Bousquet, 2006 Pterostichus barryorum Ball, 1962 Pterostichus basilobatus Sasakawa, 2005 Pterostichus basipunctatus Straneo, 1955 Pterostichus batesi (Tschitscherine, 1894) Pterostichus belizini Lutshnik, 1933 Pterostichus bellatrix (Tschitscherine, 1895) Pterostichus belousovi Kryzhanovskij, 1989 Pterostichus beneshi Sciaky, 1996 Pterostichus berezowskii (Tschitscherine, 1898) Pterostichus besucheti (Straneo, 1988) Pterostichus beyeri Vandyke, 1925 Pterostichus bhadravati Jedlicka, 1965 Pterostichus bhunetansis Davies, 2004 Pterostichus bhutanensis Morvan, 1980 Pterostichus bicolor Fischer von Waldheim, 1829 Pterostichus bielzii (Fuss, 1858) Pterostichus biexcisus Straneo, 1955 Pterostichus bifidiphallus Nemoto, 1988 Pterostichus bifoveolatus Park; Kwon & Lafer, 1996 Pterostichus binotatus Fischer von Waldheim, 1829 Pterostichus biocryus Ball, 1962 Pterostichus bischoffianus Jedlicka, 1936 Pterostichus bisetosus (Straneo, 1938) Pterostichus bisulcitarsis N.Ito, 2012 Pterostichus bituberculatus Tschitscherine, 1899 Pterostichus blandulus L.Miller, 1859 Pterostichus blodgettensis Will, 2007 Pterostichus blumenthali Heinz, 1965 Pterostichus bogatshevi (Kryzhanovskij, 1983) Pterostichus borcka Jedlicka, 1963 Pterostichus bousqueti Bergdahl, 2011 Pterostichus bowanus Sciaky, 1997 Pterostichus brachylobus Kavanaugh and LaBonte, 2006 Pterostichus brachypterus Chaudoir, 1878 Pterostichus brancuccii Straneo, 1982 Pterostichus breuili Jeannel, 1919 Pterostichus brevicornis Kirby, 1837 Pterostichus brevilama Straneo, 1989 Pterostichus brevipennis (Chevrolat, 1840) Pterostichus brevis (Duftschmid, 1812) Pterostichus breviusculus R.F. Sahlberg, 1844 Pterostichus brittoni Habu, 1958 Pterostichus bruckii Schaum, 1859 Pterostichus brunneipennis Straneo, 1955 Pterostichus bryanti (Vandyke, 1951) Pterostichus bryantoides Ball, 1962 Pterostichus bucolicus Casey, 1913 Pterostichus buglaniensis Kirschenhofer, 1981 Pterostichus bullatus Allegro & Sciaky, 2010 Pterostichus bungei (Tschitscherine, 1894) Pterostichus bureli Morvan, 1994 Pterostichus burjaticus Poppius, 1906 Pterostichus burkhan O.Berlov & Anistschenko, 2005 Pterostichus burmeisteri Heer, 1838 Pterostichus bythiniensis (Maran, 1944) Pterostichus cacumensis Ball, 1966 Pterostichus californicus (Dejean, 1828) Pterostichus caligans G.Horn, 1891 Pterostichus calvitarsis Breit, 1912 Pterostichus campania Andrewes, 1937 Pterostichus campbelli Bousquet, 1985 Pterostichus canallatus Casey, 1913 Pterostichus cancellatus (Motschulsky, 1860) Pterostichus cantabricus Schaufuss, 1862 Pterostichus cantalicus (Chaudoir, 1868) Pterostichus capitatus Chaudoir, 1850 Pterostichus capito (Tschitscherine, 1900) Pterostichus capitolinus Kurnakov, 1962 Pterostichus caribou Ball, 1962 Pterostichus carinatus Sciaky & Facchini, 2003 Pterostichus carinthiacus Meschnigg, 1946 Pterostichus carolinus Darlington, 1932 Pterostichus carradei Gautier des Cottes, 1866 Pterostichus carri Angus; Galian; Wrase & Chaladze, 2009 Pterostichus carsticus Kasahara & Y.ito, 1989 Pterostichus casalei Straneo, 1984 Pterostichus casaleianus Kirschenhofer, 1991 Pterostichus caspius (Ménétriés, 1832) Pterostichus castaneus (Dejean, 1828) Pterostichus castanipes (Ménétriés, 1843) Pterostichus castor Goulet and Bousquet, 1983 Pterostichus catalonicus J.Daniel, 1906 Pterostichus catei Sciaky & Wrase, 1997 Pterostichus cathaicus Sciaky, 1994 Pterostichus caucasicola (Tschitscherine, 1893) Pterostichus caucasicus (Ménétriés, 1832) Pterostichus caudicalis (Say, 1823) Pterostichus cavazzutii Allegro & Sciaky, 2010 Pterostichus cavicollis Straneo, 1984 Pterostichus cecchiniae Jakobson, 1907 Pterostichus cervenkai Sciaky, 1994 Pterostichus chainapaani J.Schmidt, 2005 Pterostichus chameleon Motschulsky, 1866 Pterostichus champenoisi Croissandeau, 1893 Pterostichus championi Andrewes, 1926 Pterostichus chasautianus Vysoky, 1984 Pterostichus chechcirensis Lafer, 1979 Pterostichus chefsuricus Reitter, 1896 Pterostichus chenpengi Li, 1992 Pterostichus chinkiangensis Kirschenhofer, 1991 Pterostichus chipewyan Ball, 1962 Pterostichus chogyesanus Park; Kwon & Lafer, 1996 Pterostichus chokaisanus Sasakawa, 2009 Pterostichus chotanensis (Tschitscherine, 1893) Pterostichus chujoi Habu, 1959 Pterostichus chujoiellus Jedlicka, 1962 Pterostichus chungkingi Jedlicka, 1932 Pterostichus chydaeus (Tschitscherine, 1897) Pterostichus circulosus Lindroth, 1966 Pterostichus clepsydra Sciaky & Wrase, 1997 Pterostichus cochlearis Hacker, 1968 Pterostichus cognatus (Dejean, 1831) Pterostichus coimbatoris Jedlicka, 1969